Apocalypse descends with Ghouls. Aiden York awakens the Apocalypse Tower Defense System, survives at Outpost 8. He meets Stella Shaw, North, Willow, establishes Dawn Base. He leads followers to defeat Ghoul army, purge internal strife, liberate Safe Zone. Showdown with Ghoul Empress leads to coexistence. He sacrifices himself to seal Progenitor Ghoul. He stays with the four female leads, leaving a hidden plot for the future.